How to remove these weird spots on the face of these drawers?

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I sanded these down using grits in this order: 50>80>120>220>320>400. They looked great. I wiped it down with a damp cloth. Waited for it to dry. Then I primed the wood, and as the wood was drying, and I was waiting to stain, I noticed these spots start to appear. To me they look like water spots. But there were not there before. Does anybody know what they are and how to make the wood more even?

q how to remove these weird spots on the face of these drawers
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